A pivotal role for CD40-mediated IL-6 production by dendritic cells during IL-17 induction in vivo

Georgia Perona-Wright, Stephen J. Jenkins, Richard A. O'Connor, Dimitrios Zienkiewicz, Henry J. McSorley, Rick M. Maizels, Stephen M. Anderton, Andrew S. MacDonald

    Research output: Contribution to journalArticlepeer-review

    Abstract

    The costimulatory requirements for Th17 development remain to be defined. In this study, we show that CD40-deficient animals immunized with the Gram-positive bacterium Propionibacterium acnes were specifically impaired in their ability to mount an IL-17 response, but not that of IFN-γ. The same cytokine imbalance resulted from in vivo priming with pathogenpulsed, CD40-deficient dendritic cells (DC). Engagement of CD40 on P. acnes-conditioned DC stimulated the release of IL-12, IL-23, and IL-6, of which IL-6 alone proved essential for Th17 differentiation. Compared with wild-type DC, priming with those lacking expression of CD40 resulted in reduced disease severity during experimental autoimmune encephalomyelitis, coincident with reduced IL-17 production. Our data delineate sequential requirements for DC expression of CD40 and production of IL-6 during Th17 polarization in vitro and in vivo, and reveal distinct costimulatory requirements for Th1 vs Th17 generation. Copyright © 2009 by The American Association of Immunologists, Inc.
    Original languageEnglish
    Pages (from-to)2808-2815
    Number of pages7
    JournalJournal of Immunology
    Volume182
    Issue number5
    DOIs
    Publication statusPublished - 1 Mar 2009

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