Neoselachians (Chondrichthyes, Elasmobranchii) from the Lower and lower Upper Cretaceous of north-eastern Spain

Extensive sampling of several Barremian and Albian-Cenomanian levels across the Aguilón, Oliete and Aliaga subbasins of the Iberian Basin, north-east Spain, yielded abundant material of new or so far poorly known neoselachians. The faunas consist of 16 different species, five of which represent new species and two new genera: Cantioscyllium brachyplicatum sp. nov., Platypterix venustulus gen. et sp. nov., Ptychotrygon pustulata sp. nov., Ptychotrygon striata sp. nov. and Iberotrygon plagiolophus gen. et sp. nov. In addition, teeth of Heterodontus cf. H. carerens, Lamniformes indet., Pteroscyllium sp., Scyliorhinidae indet., Rhinobatos sp., Spathobatis sp., Belemnobatis sp., Ptychotrygon geyeri, Ptychotrygon sp. and Celtipristis herreroi are described. The new family Ptychotrygonidae is defined. The localities comprise palaeoenvironments ranging from lacustrine and shallow lake to open marine settings. Neoselachians are almost completely absent from continental settings in the Barremian, as a result of prevailing freshwater conditions, but became more abundant in marine strata. The Albian-Cenomanian selachian assemblage is the most profuse and diverse of the three assemblages studied. It is dominated by small, benthic and near-coastal taxa, for instance Cantioscyllium and Ptychotrygon, and contains several new species, including an endemic batoid, Iberotrygon plagiolophus gen. et sp. nov.